### Persefone.jl - a model of agricultural landscapes and ecosystems in Europe. ### ### This file holds the code for the Eurasian Skylark (Alauda arvensis). ### skylarkhabitat = @habitat((@landcover() == grass || # settle on grass or arable land (but not maize) (@landcover() == agriculture && @cropname() != "maize")) && @distancetoedge() >= 5) # at least 50m from other habitats #XXX this ought to check for distance to forest and builtup, # but that's very expensive (see below) # @distanceto(forest) > 5 && # at least 50m from forest edges # @distanceto(builtup) > 5) # and from anthropogenic structures """ Skylark *Alauda arvensis* is a common and charismatic species of agricultural landscapes. At the moment, this implementation is still in development. **Sources:** - Bauer, H.-G., Bezzel, E., & Fiedler, W. (Eds.). (2012). Das Kompendium der Vögel Mitteleuropas: Ein umfassendes Handbuch zu Biologie, Gefährdung und Schutz (Einbändige Sonderausg. der 2., vollständig überarb. und erw. Aufl. 2005). AULA-Verlag - Glutz von Blotzheim, Urs N. (Ed.). (1985). Handbuch der Vögel Mitteleuropas. Bd. 10. Passeriformes (Teil 1) 1. Alaudidae - Hirundidae. AULA-Verlag, Wiesbaden. ISBN 3-89104-019-9 """ @species Skylark begin #XXX use Unitful.jl eggtime::Int64 = 11 # 11 days from laying to hatching eggpredationmortality::Float64 = 0.03 # per-day egg mortality from predation nestharvestmortality::Float64 = 0.9 # egg/nestling mortality after a harvest event (XXX guess) nestlingtime::Union{Int64,UnitRange{Int64}} = 7:11 # 7-11 days from hatching to leaving nest nestlingpredationmortality::Float64 = 0.03 # per-day nestling mortality from predation fledglingtime::Union{Int64,UnitRange{Int64}} = 25:30 # 25-30 days from hatching to independence fledglingharvestmortality::Float64 = 0.5 # fledgling mortality after harvest fledglingpredationmortality::Float64 = 0.01 # per-day fledgling mortality from predation firstyearmortality::Float64 = 0.38 # total mortality in the first year after independence migrationdates::Tuple = () # is defined by each individual in @create(Skylark) migrationmortality::Float64 = 0.33 # chance of dying during the winter mate::Int64 = -1 # the agent ID of the mate (-1 if none) nest::Tuple = () # coordinates of current nest nestingbegin::Tuple{Int64,Int64} = (April, 10) # begin nesting in the middle of April nestbuildingtime::Union{Int64,UnitRange{Int64}} = 4:5 # 4-5 days needed to build a nest (doubled for first nest) nestcompletion::Int64 = 0 # days left until the nest is built eggsperclutch::Union{Int64,UnitRange{Int64}} = 2:5 # 2-5 eggs laid per clutch clutch::Vector{Int64} = Vector{Int64}() # IDs of offspring in current clutch breedingdelay::Int64 = 18 # wait 18 days after hatching to start a new brood nestingend::Int64 = July # last month of nesting habitats::Function = skylarkhabitat end """ As an egg, simply check for mortality and hatching. """ @phase Skylark egg begin @kill(self.eggpredationmortality, "predation") @respond(harvesting, @kill(self.nestharvestmortality, "harvest")) if self.age == self.eggtime @setphase(nestling) end end """ As a nestling, simply check for mortality and fledging. """ @phase Skylark nestling begin #TODO add feeding & growth @kill(self.nestlingpredationmortality, "predation") @respond(harvesting, @kill(self.nestharvestmortality, "harvest")) if self.age == self.nestlingtime+self.eggtime @setphase(fledgling) end end """ As a fledgling, move around a little, but mainly wait for maturity and check mortality. """ @phase Skylark fledgling begin #TODO add feeding & growth @kill(self.fledglingpredationmortality, "predation") @walk("random") #TODO add movement following the parents if self.age == self.fledglingtime+self.eggtime @kill(self.firstyearmortality, "first year mortality") #XXX mechanistic? @setphase(nonbreeding) end end """ As a non-breeding adult, move around with other individuals and check for migration. """ @phase Skylark nonbreeding begin # flocking behaviour - follow a random neighbour or move randomly #TODO add feeding and mortality, respect habitat when moving neighbours = @neighbours(10) #XXX magic number isempty(neighbours) ? @walk("random", 5) : @follow(@rand(neighbours), 2) # check if the bird migrates leave, arrive = self.migrationdates m, d = monthday(model.date) migrate = (((m < arrive[1]) || (m == arrive[1] && d < arrive[2])) || ((m > leave[1]) || (m == leave[1] && d >= leave[2]))) if migrate #FIXME not all migrate? @kill(self.migrationmortality, "migration") returndate = Date(year(model.date)+1, arrive[1], arrive[2]) @setphase(mating) @migrate(returndate) end end """ Move around until a mate is found. """ @phase Skylark mating begin # if we've found a mate, wait for nesting begin and then go to the next phase if self.mate != -1 if !@isalive(self.mate) self.mate = -1 return end m, d = monthday(model.date) nest = ((m == self.nestingbegin[1] && d >= self.nestingbegin[2] && @chance(0.05)) || (m > self.nestingbegin[1])) #XXX why the chance? nest && @setphase(nestbuilding) return end # look for a mate among the neighbouring birds, or move randomly for n in @neighbours(50) #XXX magic number if n.sex != self.sex && n.phase == mating && n.mate == -1 self.mate = n.id n.mate = self.id @debug "$(animalid(self)) and $(animalid(n)) have mated." return end end #@debug("$(animalid(self)) didn't find a mate.") @walk("random", 10) #XXX magic number end """ Females select a location and build a nest. Males do nothing. (Sound familiar?) """ @phase Skylark nestbuilding begin if !@isalive(self.mate) self.mate = -1 @setphase(nonbreeding) return end if self.sex == female if isempty(self.nest) # try to find a nest in the neighbourhood, or move on nestlocation = @randompixel(10, self.habitats) #XXX magic number if isnothing(nestlocation) @walk("random", 20) #XXX magic number else # if we've found a location, start the clock on the building time # (building time doubles for the first nest of the year) self.nest = nestlocation self.nestcompletion = @rand(self.nestbuildingtime) month(model.date) == self.nestingbegin[1] && (self.nestcompletion *= 2) @debug "$(animalid(self)) is building a nest." end else # wait while nest is being built, then lay eggs and go to next phase if self.nestcompletion > 0 self.nestcompletion -= 1 else #XXX more accurately, a female lays one egg per day, not all at once @reproduce(@rand(self.eggsperclutch),self.mate) @setphase(breeding) end end else # males stay near the female mate = @animal(self.mate) @follow(mate, 5) mate.phase == breeding && @setphase(breeding) end end """ Do lots of foraging (not yet implemented). """ @phase Skylark breeding begin #TODO forage (move random) for offspring in self.clutch # check if offspring are still alive and juvenile, else remove from clutch if !@isalive(offspring) || @animal(offspring).phase == nonbreeding filter!(x -> x != offspring, self.clutch) end end # if all young have fledged, move to nonbreeding (if it's July) or breed again if isempty(self.clutch) self.nest = () month(model.date) >= self.nestingend ? @setphase(nonbreeding) : @setphase(nestbuilding) end end """ migrationdates(skylark, model) Select the dates on which this skylark will leave for / return from its migration, based on observed migration patterns. """ function migrationdates(skylark::Animal, model::SimulationModel) #TODO this ought to be temperature-dependent and dynamic minleave = skylark.sex == female ? (September, 15) : (October, 1) minarrive = skylark.sex == male ? (February, 15) : (March, 1) deltaleave = @rand(0:45) #XXX ought to be normally distributed deltaarrive = @rand(0:15) #XXX ought to be normally distributed leave = monthday(Date(2001, minleave[1], minleave[2]) + Day(deltaleave)) arrive = monthday(Date(2001, minarrive[1], minarrive[2]) + Day(deltaarrive)) (leave, arrive) end """ Initialise a skylark individual. Selects migration dates and checks if the bird should currently be on migration. Also sets other individual-specific variables. """ @create Skylark begin @debug "Added $(animalid(self)) at $(self.pos)" # calculate migration dates for this individual self.migrationdates = migrationdates(self, model) leave, arrive = self.migrationdates m, d = monthday(model.date) migrate = (((m < arrive[1]) || (m == arrive[1] && d < arrive[2])) || ((m > leave[1]) || (m == leave[1] && d >= leave[2]))) if migrate returndate = Date(year(model.date), arrive[1], arrive[2]) model.date != @param(core.startdate) && (returndate += Year(1)) @migrate(returndate) end # set individual life-history parameters that are defined as ranges for the species self.nestlingtime = @rand(self.nestlingtime) self.fledglingtime = @rand(self.fledglingtime) #TODO other stuff? end @populate Skylark begin habitat = skylarkhabitat initphase = mating birthphase = egg popdensity = 300 #XXX use Unitful.jl pairs = true end